┌─ Escherichia coli (42-441/446) K12 (121)  
                                                   ┌┴─ Escherichia coli (42-441/446) H299   
                                                  ┌┴─ Escherichia coli (42-441/446) O157:H7 EDL933 (1)  
                                                 ┌┴─ Escherichia coli (3-390/395) B088 (3)  
     ┌───────────────────────────────────────────┴─ Escherichia coli (42-441/446) UTI89 (1)  
┌────┴────────────────────────────────────────────── Escherichia coli (37-438/438) MS 69-1 (2) [2] 
┤                                             ┌── Escherichia coli (40-432/432) M718 (4)  
│               ┌─────────────────────────────┴── fig|6666666.5357.peg.2015 TY-2482 (40-432/432) strain (2)  
└───────────────┤                                                   ┌─ Escherichia coli (42-442/444) MS 69-1 (2)  
                │                  ┌────────────────────────────────┴─ Escherichia coli (42-442/444) MS 107-1   
                └──────────────────┤            ┌─ Escherichia coli (32-433/433) E24377A   
                                   └────────────┤┌─ fig|1040638.4.peg.836 O104:H4 str. LB226692 (8)  
                                                └┤┌─ Escherichia coli (32-433/433) K12 (24)  
                                                 └┤ ┌─ Escherichia coli DH1   
                                                  └┬┴─ Escherichia coli O103:H2 str. 12009   
                                                   └┬─ Escherichia coli 042 (91)  
                                                    └─ Escherichia coli O157:H7 EDL933   
  

  
C4-dicarboxylate transporter DcuA
 (5)
  
C4-dicarboxylate transporter DcuB
 (5)
  
Anaerobic C4-dicarboxylate transporter DcuB
 (2)
  
Anaerobic C4-dicarboxylate transporter dcuA
 (1)
  
anaerobic C4-dicarboxylate membrane transporter
 (1)
  
Anaerobic C4-dicarboxylate transporter
 (1)
  
 Others 
(2)