┌─────────────────────────────────── Escherichia coli (385-442/446) K12 (133)  
┌───────────┴──────────────────────────────────────────────────────── Escherichia coli (383-438/438) MS 69-1 (2) [2] 
┤                           ┌───── Escherichia coli (372-432/432) M718 (4)  
└─┬─────────────────────────┴────── fig|6666666.5357.peg.2015 TY-2482 (372-432/432) strain (2) [2] 
  └──────────────────────────────┬──────────────────── Escherichia coli (386-443/444) MS 69-1 (3)  
                                 └──────────────┬─ fig|1040638.4.peg.836 O104:H4 (343-402/402) str. LB226692 (9)  
                                                └─┬── Escherichia coli (193-252/252) OP50   
                                                  └─ Escherichia coli (374-433/433) K12 (121)  
  

  
C4-dicarboxylate transporter DcuA
 (2)
  
C4-dicarboxylate transporter DcuB
 (1)
  
Anaerobic C4-dicarboxylate transporter DcuB
 (1)
  
anaerobic C4-dicarboxylate membrane transporter
 (1)
  
Anaerobic C4-dicarboxylate transporter
 (1)
  
 Others 
(2)